The Wonderful Precure! anime is set in Animal Town, a town where animals and people live together in harmony. Second-year junior high school student Iroha Inukai is best friends with her dog named Komugi. Out for a walk one day, Iroha and Komugi encounter a mysterious creature named Garugaru, who goes on a rampage in the city. Komugi suddenly transforms into a Pretty Cure named Cure Wonderful, to protect Iroha.